Pseudoscorpiones, False Scorpions: Chthoniidae Daday, 1889 accepted

WAC:	urn:lsid:wac.nmbe.ch:name:b2ab88ef-50c8-4984-828d-256483030502
ITIS:	urn:lsid:itis.gov:itis_tsn:746809
GBIF:	7652
Fauna Europaea:	082a4bc9-5f15-402f-ac33-05e3c3f759b5

Species in this family

13 genera, 343 species:

Cantabrochthonius

Cantabrochthonius asturiensis (Beier, 1955) accepted

Chthonius

Chthonius absoloni Beier, 1938 accepted

Chthonius agazzii Beier, 1966 accepted

Chthonius alpicola Beier, 1951 accepted

Chthonius apollinis Mahnert, 1978 accepted

Chthonius aquasanctae Ćurčić & Rađa, 2011 accepted

Chthonius azerbaidzhanus Schawaller & Dashdamirov, 1988 accepted

Chthonius balazuci Vachon, 1963 accepted

Chthonius bogovinae Ćurčić, 1972 accepted

Chthonius campaneti Zaragoza & Vadell, 2013 accepted

Chthonius caoduroi Callaini, 1987 accepted

Chthonius caprai Gardini, 1977 accepted

Chthonius carinthiacus Beier, 1951 accepted

Chthonius cavernarum Ellingsen, 1909 accepted

Chthonius cavophilus Hadži, 1939 accepted

Chthonius cebenicus Leclerc, 1981 accepted

Chthonius cephalotes (Simon, 1875) accepted

Chthonius chamberlini (Leclerc, 1983) accepted

Chthonius comottii Inzaghi, 1987 accepted

Chthonius croaticus Ćurčić & Rađa, 2012 accepted

Chthonius cryptus Chamberlin, 1962 accepted

Chthonius dacnodes Navás, 1918 accepted

Chthonius decoui Georgescu & Cǎpuşe, 1994 accepted

Chthonius delmastroi Gardini, 2009 accepted

Chthonius densedentatus Beier, 1938 accepted

Chthonius doderoi Beier, 1930 accepted

Chthonius elongatus Lazzeroni, 1970 accepted

Chthonius euganeus Gardini, 1991 accepted

Chthonius exarmatus Beier, 1939 accepted

Chthonius gallettii Gardini, 2021 accepted

Chthonius gentianae Gardini, 2021 accepted

Chthonius gjirokastri Ćurčić, Rađa & Dimitrijević, 2008 accepted

Chthonius globocicae Ćurčić, Dimitrijević & Tomić, 2008 accepted

Chthonius graecus Beier, 1963 accepted

Chthonius guglielmii Callaini, 1986 accepted

Chthonius halberti Kew, 1916 accepted

Chthonius herbarii Mahnert, 1980 accepted

Chthonius herminii Gardini, 2021 accepted

Chthonius heterodactylus Tömösváry, 1883 accepted

Chthonius heurtaultae Leclerc, 1981 accepted

Chthonius horridus Beier, 1934 accepted

Chthonius hungaricus Mahnert, 1981 accepted

Chthonius ilvensis Beier, 1963 accepted

Chthonius imperator Mahnert, 1978 accepted

Chthonius inguscioi Gardini, 2021 accepted

Chthonius ischnocheles (Hermann, 1804) accepted

Chthonius ischnocheles ischnocheles (Hermann, 1804) accepted

Chthonius ischnocheloides Beier, 1973 accepted

Chthonius italicus Beier, 1930 accepted

Chthonius iugoslavicus Ćurčić, 1972 accepted

Chthonius jonicus Beier, 1931 accepted

Chthonius jugorum Beier, 1952 accepted

Chthonius kladanjensis Ćurčić, Dimitrijević and Rađa, 2011 accepted

Chthonius kosovensis Ćurčić, 2011 accepted

Chthonius lanai Gardini, 2021 accepted

Chthonius lanzai Caporiacco, 1947 accepted

Chthonius lanzai lanzai Caporiacco, 1947 accepted

Chthonius lanzai vannii Callaini, 1968 accepted

Chthonius latidentatus Ćurčić, 1972 accepted

Chthonius leoi (Callaini, 1988) accepted

Chthonius lesnik Ćurčić, 1994 accepted

Chthonius lessiniensis Schawaller, 1982 accepted

Chthonius ligusticus Beier, 1930 accepted

Chthonius ligusticus brevis Cîrdei, Bulimar & Malcoci, 1967 accepted

Chthonius lindbergi Beier, 1956 accepted

Chthonius lucifugus Mahnert, 1977 accepted

Chthonius lupinus Ćurčić, Dimitrijević & Rađa, 2011 accepted

Chthonius macedonicus Ćurčić, 1972 accepted

Chthonius magnificus Beier, 1938 accepted

Chthonius makirina Ćurčić & Rađa, 2012 accepted

Chthonius marciai Gardini, 2021 accepted

Chthonius mauritanicus (Callaini, 1988) accepted

Chthonius mayi Heurtault-Rossi, 1968 accepted

Chthonius mazaurici Leclerc, 1981 accepted

Chthonius mazaurici coironi Leclerc, 1981 accepted

Chthonius mazaurici mazaurici Leclerc, 1981 accepted

Chthonius microphthalmus Simon, 1879 accepted

Chthonius minotaurus Henderickx, 1997 accepted

Chthonius monicae Boghean, 1989 accepted

Chthonius motasi Dumitresco & Orghidan, 1964 accepted

Chthonius multidentatus Beier, 1963 accepted

Chthonius nicolosii Gardini, 2021 accepted

Chthonius occultus Beier, 1939 accepted

Chthonius ognjankae Ćurčić, 1997 accepted

Chthonius ohridanus Ćurčić, 1997 accepted

Chthonius onaei Ćurčić, Dimitrijević, Rađa, Ćurčić & Milinčić, 2010 accepted

Chthonius orthodactyloides Beier, 1967 accepted

Chthonius orthodactylus (Leach, 1817) accepted

Chthonius pagus Ćurčić & Rađa, 2012 accepted

Chthonius paolettii Beier, 1973 accepted

Chthonius persimilis Beier, 1939 accepted

Chthonius petrochilosi Heurtault, 1972 accepted

Chthonius ponticoides Mahnert, 1975 accepted

Chthonius ponticus Beier, 1965 accepted

Chthonius porevidi Ćurčić, Makarov & Lučić, 1998 accepted

Chthonius pristani Ćurčić, 2011 accepted

Chthonius protobosniacus Ćurčić, Dimitrijević & Rađa, 2011 accepted

Chthonius prove Ćurčić, Dimitrijević & Makarov, 1997 accepted

Chthonius pusillus Beier, 1947 accepted

Chthonius pygmaeus Beier, 1934 accepted

Chthonius radigost Ćurčić, 1997 accepted

Chthonius radjai Ćurčić, 1988 accepted

Chthonius raridentatus Hadži, 1930 accepted

Chthonius reductus Beier, 1939 accepted

Chthonius ressli Beier, 1956 accepted

Chthonius sestasi Mahnert, 1980 accepted

Chthonius shelkovnikovi Redikorzev, 1930 accepted

Chthonius stammeri Beier, 1942 accepted

Chthonius stevanovici Ćurčić, 1986 accepted

Chthonius strinatii Mahnert, 1975 accepted

Chthonius submontanus Beier, 1963 accepted

Chthonius subterraneus Beier, 1931 accepted

Chthonius subterraneus meuseli Beier, 1939 accepted

Chthonius subterraneus subterraneus Beier, 1931 accepted

Chthonius tenuis L. Koch, 1873 accepted

Chthonius thessalus Mahnert, 1980 accepted

Chthonius torakensis Ćurčić & Rađa, 2010 accepted

Chthonius trebinjensis Beier, 1938 accepted

Chthonius troglobius Hadži, 1937 accepted

Chthonius troglodites Redikorzev, 1928 accepted

Chthonius tzanoudakisi Mahnert, 1975 accepted

Chthonius vodan Ćurčić, 1997 accepted

Chthonius zmaj Ćurčić, 1997 accepted

Ephippiochthonius

Ephippiochthonius aeneae (Gardini, 2013) accepted

Ephippiochthonius aguileraorum (Carabajal Márquez, García Carrillo & Rodríguez Fernández, 2000) accepted

Ephippiochthonius aini Zaragoza, 2017 accepted

Ephippiochthonius altamurae (Gardini, 2013) accepted

Ephippiochthonius amatei (Carabajal Márquez, García Carrillo & Rodríguez Fernández, 2001) accepted

Ephippiochthonius anatolicus (Beier, 1969) accepted

Ephippiochthonius andalucia Zaragoza, 2017 accepted

Ephippiochthonius anophthalmus (Ellingsen, 1908) accepted

Ephippiochthonius apulicus (Beier, 1958) accepted

Ephippiochthonius atlantis (Mahnert, 1980) accepted

Ephippiochthonius aurouxi Zaragoza, 2017 accepted

Ephippiochthonius balearicus (Mahnert, 1977) accepted

Ephippiochthonius bellesi (Mahnert, 1989) accepted

Ephippiochthonius benimaquia Zaragoza, 2017 accepted

Ephippiochthonius bidentatus (Beier, 1938) accepted

Ephippiochthonius boldorii (Beier, 1934) accepted

Ephippiochthonius bolivari (Beier, 1930) accepted

Ephippiochthonius borissketi (Ćurčić & Sarbu, 2014) accepted

Ephippiochthonius cabreriensis (Mahnert, 1993) accepted

Ephippiochthonius caceresi Zaragoza, 2017 accepted

Ephippiochthonius castellonensis Zaragoza, 2017 accepted

Ephippiochthonius catalonicus (Beier, 1939) accepted

Ephippiochthonius chius (Schawaller, 1990) accepted

Ephippiochthonius cikolae (Ćurčić & Rađa, 2010) accepted

Ephippiochthonius civitatisveti (Ćurčić & Rađa, 2011) accepted

Ephippiochthonius comasi Zaragoza, 2017 accepted

Ephippiochthonius concii (Beier, 1953) accepted

Ephippiochthonius corcyraeus (Mahnert, 1976) accepted

Ephippiochthonius corsicus (Callaini, 1981) accepted

Ephippiochthonius creticus (Mahnert, 1980) accepted

Ephippiochthonius daedaleus (Mahnert, 1980) accepted

Ephippiochthonius distinguendus (Beier, 1930) accepted

Ephippiochthonius elymus (Gardini, 2013) accepted

Ephippiochthonius etruscus (Gardini, 2013) accepted

Ephippiochthonius fadriquei Zaragoza, 2017 accepted

Ephippiochthonius fuscimanus (Simon, 1900) accepted

Ephippiochthonius galcerani Zaragoza, 2017 accepted

Ephippiochthonius gallii (Gardini, 2013) accepted

Ephippiochthonius gasparoi (Gardini, 1989) accepted

Ephippiochthonius genuensis (Gardini, 1990) accepted

Ephippiochthonius gestroi (Simon, 1896) accepted

Ephippiochthonius gibbus (Beier, 1953) accepted

Ephippiochthonius girgentiensis (Mahnert, 1982) accepted

Ephippiochthonius giustii (Callaini, 1981) accepted

Ephippiochthonius gonzalezi Zaragoza, 2017 accepted

Ephippiochthonius grafittii (Gardini, 1981) accepted

Ephippiochthonius henderickxi Zaragoza, 2017 accepted

Ephippiochthonius hiberus (Beier, 1930) accepted

Ephippiochthonius hispanus (Beier, 1930) accepted

Ephippiochthonius ibiza Zaragoza, 2017 accepted

Ephippiochthonius insularis (Beier, 1938) accepted

Ephippiochthonius intemelius (Gardini, 2013) accepted

Ephippiochthonius kabylicus (Callaini, 1983) accepted

Ephippiochthonius kemza (Ćurčić, Lee & Makarov, 1993) accepted

Ephippiochthonius kewi (Gabbutt, 1966) accepted

Ephippiochthonius kupalo (Ćurčić, 1997) accepted

Ephippiochthonius lagadini (Ćurčić & Rađa, 2011) accepted

Ephippiochthonius latellai (Gardini, 2013) accepted

Ephippiochthonius ligur (Gardini, 2013) accepted

Ephippiochthonius longesetosus (Mahnert, 1976) accepted

Ephippiochthonius lucanus (Callaini, 1984) accepted

Ephippiochthonius lychnidis (Ćurčić, 1997) accepted

Ephippiochthonius magrinii (Gardini, 2013) accepted

Ephippiochthonius mahnerti (Zaragoza, 1984) accepted

Ephippiochthonius mariolae (Carabajal Márquez, García Carrillo & Rodríguez Fernández, 2001) accepted

Ephippiochthonius maroccanus (Mahnert, 1980) accepted

Ephippiochthonius masoae Zaragoza, 2017 accepted

Ephippiochthonius mayorali (Carabajal Márquez, García Carrillo & Rodríguez Fernández, 2001) accepted

Ephippiochthonius messapicus (Gardini, 2013) accepted

Ephippiochthonius metohicus (Ćurčić, 2011) accepted

Ephippiochthonius microtuberculatus (Hadži, 1937) accepted

Ephippiochthonius minous (Mahnert, 1980) accepted

Ephippiochthonius minous minous (Mahnert, 1980) accepted

Ephippiochthonius minous peramae (Mahnert, 1980) accepted

Ephippiochthonius monguzzii (Gardini, 2013) accepted

Ephippiochthonius nanus (Beier, 1953) accepted

Ephippiochthonius nerjaensis (Carabajal Márquez, García Carrillo & Rodríguez Fernández, 2001) accepted

Ephippiochthonius nidicola (Mahnert, 1979) accepted

Ephippiochthonius nudipes (Mahnert, 1982) accepted

Ephippiochthonius oryzis Kolesnikov, Turbanov & Gongalsky, 2019 accepted

Ephippiochthonius pieltaini (Beier, 1930) accepted

Ephippiochthonius platakisi (Mahnert, 1980) accepted

Ephippiochthonius pliginskyi Turbanov & Kolesnikov, 2021 accepted

Ephippiochthonius ponsi (Mahnert, 1993) accepted

Ephippiochthonius portugalensis Zaragoza, 2017 accepted

Ephippiochthonius poseidonis (Gardini, 1993) accepted

Ephippiochthonius pyrenaicus (Beier, 1934) accepted

Ephippiochthonius remyi (Heurtault, 1975) accepted

Ephippiochthonius rhizon (Ćurčić, 2013) accepted

Ephippiochthonius riberai Zaragoza, 2017 accepted

Ephippiochthonius rimicola (Mahnert, 1993) accepted

Ephippiochthonius rogoi (Ćurčić, Rađa & Dimitrijević, 2008) accepted

Ephippiochthonius romanicus (Beier, 1935) accepted

Ephippiochthonius samius (Mahnert, 1982) accepted

Ephippiochthonius sardous (Gardini, 2008) accepted

Ephippiochthonius sarmaticus Kolesnikov, Turbanov & Gongalsky, 2019 accepted

Ephippiochthonius schmalfussi (Schawaller, 1990) accepted

Ephippiochthonius scythicus (Georgescu & Cǎpuşe, 1994) accepted

Ephippiochthonius serbicus (Hadži, 1937) accepted

Ephippiochthonius serengei Zaragoza, 2017 accepted

Ephippiochthonius sevai Zaragoza, 2017 accepted

Ephippiochthonius siculus (Beier, 1961) accepted

Ephippiochthonius siscoensis (Heurtault, 1975) accepted

Ephippiochthonius sulphureus (Gardini, 2013) accepted

Ephippiochthonius tarraconensis Zaragoza, 2017 accepted

Ephippiochthonius tauroscythicus Turbanov & Kolesnikov, 2021 accepted

Ephippiochthonius tetrachelatus (Preyssler, 1790) accepted

Ephippiochthonius timacensis (Ćurčić, Dimitrijević & Stojanović, 2012) accepted

Ephippiochthonius troglophilus (Beier, 1930) accepted

Ephippiochthonius tuberculatus (Hadži, 1937) accepted

Ephippiochthonius tyrrhenicus (Gardini, 2013) accepted

Ephippiochthonius vachoni (Heurtault-Rossi, 1963) accepted

Ephippiochthonius vicenae Zaragoza, 2017 accepted

Ephippiochthonius vid (Ćurčić, 1997) accepted

Ephippiochthonius volkeri Turbanov & Kolesnikov, 2021 accepted

Ephippiochthonius zaballosi Zaragoza, 2017 accepted

Ephippiochthonius zoiai (Gardini, 1990) accepted

Globochthonius

Globochthonius abnormis (Beier, 1939) accepted

Globochthonius argirocastronis (Ćurčić, Rađa & Dimitrijević, 2007) accepted

Globochthonius caligatus (Beier, 1938) accepted

Globochthonius cavernicola (Beier, 1938) accepted

Globochthonius cerberus (Beier, 1938) accepted

Globochthonius daorsoni (Ćurčić & Rađa, 2014) accepted

Globochthonius globifer (Simon, 1879) accepted

Globochthonius globimanus (Ćurčić & Rađa, 2011) accepted

Globochthonius longimanus (Ćurčić & Rađa, 2011) accepted

Globochthonius medeonis (Ćurčić, 2011) accepted

Globochthonius montis (Ćurčić, 2013) accepted

Globochthonius pancici (Ćurčić, 1972) accepted

Globochthonius perun (Ćurčić, 1997) accepted

Globochthonius petroupauli (Ćurčić & Rađa, 2011) accepted

Globochthonius poeninus (Mahnert, 1979) accepted

Globochthonius polychaetus (Hadži, 1937) accepted

Globochthonius purgo (Ćurčić, Lee & Makarov, 1993) accepted

Globochthonius satapliaensis (Schawaller & Dashdamirov, 1988) accepted

Globochthonius simplex (Beier, 1939) accepted

Globochthonius spelaeophilus (Hadži, 1930) accepted

Globochthonius vandeli (Dumitresco & Orghidan, 1964) accepted

Lechytia

Lechytia anatolica Beier, 1965 accepted

Microchthonius

Microchthonius babasarka Ćurčić, Rađa & Dimitrijević, 2015 accepted

Microchthonius dernisi Ćurčić & Rađa, 2012 accepted

Microchthonius elegantissimus B. P. M. Ćurčić, Rađa, S. B. Ćurčić, Ilić, Tomić & Makarov, 2013 accepted

Microchthonius karamani (Hadži, 1933) accepted

Microchthonius kasteli Ćurčić & Rađa, 2014 accepted

Microchthonius lorko Ćurčić, Rađa & Dimitrijević, 2015 accepted

Microchthonius rogatus (Beier, 1938) accepted

Microchthonius solentanus Ćurčić & Rađa, 2013 accepted

Microchthonius tragurion Ćurčić & Rađa, 2013 accepted

Mundochthonius

Mundochthonius alpinus Beier, 1947 accepted

Mundochthonius basarukini Schawaller, 1989 accepted

Mundochthonius carpaticus Rafalski, 1948 accepted

Mundochthonius decoui Dumitresco & Orghidan, 1970 accepted

Mundochthonius gallaecicus Zaragoza & Harvey, 2006 accepted

Mundochthonius styriacus Beier, 1971 accepted

Mundochthonius ussuricus Beier, 1979 accepted

Neobalkanella

Neobalkanella psoglavi (Ćurčić, 1990) accepted

Occidenchthonius

Occidenchthonius alandroalensis Zaragoza & Reboleira, 2018 accepted

Occidenchthonius algharbicus Zaragoza & Reboleira, 2018 accepted

Occidenchthonius ambrosiae (Carabajal Márquez, García Carrillo & Rodríguez Fernández, 2012) accepted

Occidenchthonius anae Zaragoza, 2017 accepted

Occidenchthonius beieri Zaragoza, 2017 accepted

Occidenchthonius berninii (Callaini, 1983) accepted

Occidenchthonius bullonorum (Carabajal Márquez, García Carrillo & Rodríguez Fernández, 2012) accepted

Occidenchthonius canariensis (Beier, 1965) accepted

Occidenchthonius cardosoi (Zaragoza, 2012) accepted

Occidenchthonius cassolai (Beier, 1973) accepted

Occidenchthonius cazorlensis (Carabajal Márquez, García Carrillo & Rodríguez Fernández, 2001) accepted

Occidenchthonius dubius (Mahnert, 1993) accepted

Occidenchthonius duecensis Zaragoza & Reboleira, 2018 accepted

Occidenchthonius ebo Zaragoza, 2017 accepted

Occidenchthonius espanyoli (Zaragoza & Pérez, 2013) accepted

Occidenchthonius felgueraorum Zaragoza, 2017 accepted

Occidenchthonius gardinii Zaragoza, 2017 accepted

Occidenchthonius giennensis (Zaragoza & Pérez, 2013) accepted

Occidenchthonius goncalvesi Zaragoza & Reboleira, 2018 accepted

Occidenchthonius gracilimanus (Mahnert, 1997) accepted

Occidenchthonius hoerwegi Zaragoza, 2017 accepted

Occidenchthonius lencinai Zaragoza, 2017 accepted

Occidenchthonius machadoi (Vachon, 1940) accepted

Occidenchthonius mateui Zaragoza, 2017 accepted

Occidenchthonius minutus (Vachon, 1940) accepted

Occidenchthonius montagudi Zaragoza, 2017 accepted

Occidenchthonius morenoi (Carabajal Márquez, García Carrillo & Rodríguez Fernández, 2011) accepted

Occidenchthonius murcia Zaragoza, 2017 accepted

Occidenchthonius ortunoi Zaragoza, 2017 accepted

Occidenchthonius parmensis (Beier, 1963) accepted

Occidenchthonius perezi (Carabajal Márquez, García Carrillo & Rodríguez Fernández, 2011) accepted

Occidenchthonius pinai (Zaragoza, 1985) accepted

Occidenchthonius riopar Zaragoza, 2017 accepted

Occidenchthonius ruizporteroae (Carabajal Márquez, García Carrillo & Rodríguez Fernández, 2001) accepted

Occidenchthonius sendrai (Zaragoza, 1985) accepted

Occidenchthonius serranoi Zaragoza, 2017 accepted

Occidenchthonius setosus (Mahnert, 1993) accepted

Occidenchthonius thaleri (Gardini, 2009) accepted

Occidenchthonius torremarinae (Carabajal Márquez, García Carrillo & Rodríguez Fernández, 2012) accepted

Occidenchthonius vachoni Zaragoza & Reboleira, 2018 accepted

Occidenchthonius ventalloi (Beier, 1939) accepted

Occidenchthonius verai (Zaragoza, 1985) accepted

Occidenchthonius villacarrillo (Zaragoza & Pérez, 2013) accepted

Paraliochthonius

Paraliochthonius barrancoi (Carabajal Márquez, García Carrillo & Rodríguez Fernández, 2001) accepted

Paraliochthonius canariensis Vachon, 1961 accepted

Paraliochthonius cavalensis Zaragoza, 2004 accepted

Paraliochthonius curvidigitatus Mahnert, 1997 accepted

Paraliochthonius hoestlandti Vachon, 1960 accepted

Paraliochthonius martini Mahnert, 1989 accepted

Paraliochthonius mirus Mahnert, 2002 accepted

Paraliochthonius setiger (Mahnert, 1997) accepted

Paraliochthonius singularis (Menozzi, 1924) accepted

Paraliochthonius superstes (Mahnert, 1986) accepted

Paraliochthonius tenebrarum Mahnert, 1989 accepted

Spelyngochthonius

Spelyngochthonius beieri Gardini, 1994 accepted

Spelyngochthonius dewaelei Gardini, 2008 accepted

Spelyngochthonius grafittii Gardini, 1994 accepted

Spelyngochthonius heurtaultae Vachon, 1967 accepted

Spelyngochthonius provincialis Vachon & Heurtault-Rossi, 1964 accepted

Spelyngochthonius sardous Beier, 1955 accepted

Troglochthonius

Troglochthonius doratodactylus Helversen, 1968 accepted

Troglochthonius mirabilis Beier, 1939 accepted

Zaragozachthonius

Zaragozachthonius karamanianus (Hadži, 1937) accepted

Zaragozachthonius siculus Gardini, 2020 accepted